The evolution and adaptation of infanticide and siblicide

The evolution and adaptation of infanticide and siblicide INTRODUCTION LIFE HISTORY The term life history describes an organisms strategy of allocating time and energy between growth, reproduction and survival (Levin 2009; Bergon, Townsend and Harper 2008). Life history traits make up an organisms life history and include growth patterns, size at birth, size and age at sexual maturity, the number, size and sex ratio of offspring, parental care and length of life (Levin 2009; Bergon, Townsend and Harper 2008). The effect of finite resource availability on organisms for potential investment into growth, reproduction, and survival, is to set limits on life history traits, often referred to as trade-offs (Levin 2009; Bergon, Townsend and Harper 2008; Cotgreave and Forseth 2008). If an organism commits more time and energy into one specific life history trait, it will come at a cost, a reduction of time and energy that may have been available to one or more other life history traits (Levin 2009; Bergon, Townsend and Harper 2008). An example of this often observed in rela tion to reproductive strategies (Forbes and Mock 2000). Many reproductive strategies have evolved but are always subject to tradeoffs; it is a case of quantity versus quality (Forbes and Mock 2000). Life history traits respond like any other phenotypic trait to natural selection and therefore represent adaptations that have been made by organisms to their environments (Levin 2009; Cotgreave and Forseth 2008). The question; Offspring are difficult to produce and are a critical determinate of fitness. So how does siblicide and infanticide evolve as a life history strategy and are they adaptive? SIBLICIDE The demise of an individual as a result of another directly related individuals actions is referred to as siblicide (Anderson 1995; Godfray and Harper 1990; Anderson 1989). Siblicide may transpire among siblings or be brought about by the actions of parents (Hausfater and Hrdy 2008; Anderson 1995). Siblicide has been documented occurring in plants, fish, insects and mammals but has been best observed in avians (Holcomb 2001; Mock and Parker 1997). There are many hypotheses surrounding siblicide and why it occurs (Godfray and Harper 1990). Siblicide may take place as a direct form of assault on another sibling through physical attacks or expulsion from the nest, or may transpire indirectly in the form of exclusion from food by competition which results in starvation (Hausfater and Hrdy 2008). There are two forms of siblicide known as obligate and facultative siblicide (Hausfater and Hrdy 2008; Anderson 1995; Anderson 1988). The Brood Reduction hypothesis best supports facultative siblicide as an adaptive reproductive strategy that benefits both parents and surviving offspring in response to the event of unpredictable resource shortages (Forbes and Mock 2000; Mock and Parker 1997). The strategy is to hatch as many offspring as would normally be expected if conditions were optimal (Hausfater and Hrdy 2008; Forbes and Mock 2000; Anderson 1995). In the event that environmental conditions are severe and the resources available are unable to meet the demands for all of the offspring to survive, sibling rivalry ensues and is anticipated to become fatal resulting in brood reduction (Hausfater and Hrdy 2008; Forbes and Mock 2000; Mock and Parker 1997). The reduction in brood size will then allow the parents to successfully raise the remaining nestlings (Hausfater and Hrdy 2008; Forbes and Mock 2000; Godfray and Harper 1990). Blue-footed boobies are well known for facultative siblicide (Anderson 1995; Anderson 19 88). They are capable of laying between one and four eggs, the average clutch size though is usually two (Anderson 1995). When resources are plentiful all of the chicks are hatched, and fledge, however, should resources be scarce, the first hatched chick will dispatch the younger siblings (Anderson 1995). Obligate siblicide is the occurrence of siblicide regardless of resource abundance (Anderson 1989). Parents regularly produce more offspring than they can successfully fledge and it is the case that one the first born nestling will eliminate the second nestling soon after hatching (Hausfater and Hrdy 2008; Forbes and Mock 2000; Anderson 1989). The hypothesis that best supports obligate siblicide is the Insurance Egg hypothesis (Hausfater and Hrdy 2008; Forbes and Mock 2000; Anderson 1989). The Insurance Egg hypothesis is the theory that parents actually produce a second egg specifically as a backup, in the event that the first egg fails or succumbs to predation (Hausfater and Hrdy 2008; Forbes and Mock 2000; Anderson 1989). The cost of the second egg is essentially less to the parents than the benefit of producing that insurance egg should the first egg fail (Forbes and Mock 2000; Anderson 1989). This overproduction is an adaptive response to the insecurity of offspring survival or v iability (Forbes and Mock 2000; Anderson 1989. In general, very few species of bird commit obligate siblicide (Anderson 1989). The masked boobies and brown boobies are two obligately siblicidal species (Anderson 1995; Anderson 1989). Blue-footed boobies, masked boobies and brown boobies all utilize the same life history trait of asynchronous hatching (Anderson 1995; Anderson 1988). There are differences however, between the asynchronous hatching times as a result of facultative or obligative siblicide (Anderson 1995; Anderson 1988). The length of asynchronous hatching is shorter in the blue-footed boobies than the masked or brown boobies, mainly due to the form of siblicide (Anderson 1988). The effect of asynchronous hatching on the nestlings is conferred in age, size and hierarchy of the first hatched nestling over the second or third nestling and therefore, the offspring hatched first has the competitive advantage over the later hatched offspring as a result (Anderson 1988). In regard to facultative siblicide, this manipulation by the parents in staggering hatching times can be viewed as a strategy to counteract the uncertainty of resource availability by bestowing the competitive advantage on the first hatched nestling should brood reduction become necessary in the event of a shortage of resources (Hausfater and Hrdy 2008; Anderson 1988). In regard to obligative siblicide, the first hatched chick will inevitably commit siblicide and therefore the advantages of being first born come into play (Hausfater and Hrdy 2008; Anderson 1989). INFANTICIDE The term infanticide can be described as the killing of dependent offspring by individuals belonging to the same species (Hausfater and Hrdy 2008; Hiraiwa-Hasegawa 1988). Infanticide is not limited to the killing of unweaned offspring, it can occur during the reproductive cycle, for example re-absorption of the embryo or abortion, and can be committed by females and males as well as offspring or other members within the social group (Hausfater and Hrdy 2008). Infanticide has been observed in mammals, including several primate species and lions (Hausfater and Hrdy 2008; Hiraiwa-Hasegawa 1988; Packer and Pusey 1983). The act of infanticide is an adaptive behaviour strategy to enhance individual fitness (Hausfater and Hrdy 2008; Agrell and Wolff 1998; Hiraiwa-Hasegawa 1988; Packer and Pusey 1983). Infanticide of unrelated infants committed by males due to reproductive competition is supported by the sexual selection hypothesis (Kappeler and van Schaik 2004; Borries et al. 1999; Agrell a nd Wolff 1998; Hiraiwa-Hasegawa 1988). Under this hypothesis, infanticidal males secure mating opportunities and increase their chance of siring infants and therefore gain a reproductive advantage and increase fitness (Borries et al. 1999). African lions (Panthera leo) and hanuman langurs (Presbytis entellus) both live in groups consisting of one dominant male and a number of females (Hiraiwa-Hasegawa 1988). The residence time of the dominant male is usually short, approximately two years (Hiraiwa-Hasegawa 1988). Infanticide occurs as a consequence of a group takeover, one male gaining control of anothers group (Hausfater and Hrdy 2008; Kappeler and van Schaik 2004). As a result of short term dominance status of males in these instances, it is to the advantage of the usurping male to dispatch of infants within the group so he can take full advantage of the females reproductive career (Kappeler and van Schaik 2004; Borries et al. 1999). Both female langurs and lions share a common life history trait, they are almost always ready to resume sexual activity and begin reproducing after the loss of their unweaned infant; much earlier than they otherwise would if they still had care of their dependant offspring (Hausfater and Hrdy 2008; Kappeler and van Schaik 2004). There is evidence to show that female primates and lions swiftly revert to estrus after the loss of their unweaned infant (Hausfater and Hrdy 2008; Kappeler and van Schaik 2004; Borries et al. 1999; Packer and Pusey 1983). Whilst females are lactating, they are effectively unresponsive to further reproduction (Hausfater and Hrdy 2008; Packer and Pusey 1983). Therefore, the act of infanticide and the quick return of estrus as a result, ensures the females bear infants to the usurper much sooner than if the females had surviving infants (Hausfater and Hrdy 2008; Kappeler and van Schaik 2004; Borries et al. 1999; Agrell and Wolff 1998). Infanticide is undoubtedly a major disadvantage for female reproductive success even though it may well be an adaptive behavioural strategy for male reproductive success (Hausfater and Hrdy 2008; Kappeler and van Schaik 2004; Packer and Pusey 1983). As such females have developed counter strategies in an attempt to decrease their reproductive losses as a result of infanticidal males after a takeover has occurred (Hausfater and Hrdy 2008; Kappeler and van Schaik 2004; Agrell and Wolff 1998; Packer and Pusey 1983). Research into these behavioural strategies has revealed a range of different tactics employed by females in an effort to rescue their infants from almost certain death (Hausfater and Hrdy 2008; Kappeler and van Schaik 2004; Agrell and Wolff 1998; Packer and Pusey 1983). Some of the strategies employed may include leaving the group, sometimes in the company of the deposed male, aggressively defending the infant from attacks by the new male, repeatedly mating with the new male to confuse paternity of already pregnant females at the time of takeover, pseudo-estrus of already pregnant females or in some cases, the termination of an early pregnancy to avoid the inevitable (Agrell and Wolff 1998; Packer and Pusey 1983). CONCLUSION How and why siblicide and infanticide life history strategies have evolved has been the subject of great debate for many years. The results of studies conducted provide quite a lot of evidence in support of the different theories for both siblicide and infanticide being adaptive life history strategies (Kappeler and van Schaik 2004; Forbes and Mock 2000; Borries et al. 1999; Mock and Parker 1997; Agrell and Wolff 1998; Hiraiwa-Hasegawa 1988). In consideration of siblicide, studies conducted provide evidence in support of theories that facultative siblicide has evolved in response to the unreliability of resources and, obligate siblicide has evolved due to the uncertainty of survival or viability of offspring. Where infanticide is concerned, evidence favours the theory that evolution of this life history strategy is predominately an adaptive male behavioural strategy to increase reproductive success in response to short term dominant male status within groups. The act of infanticide i ncreases the males chance of successfully siring offspring while they hold the dominant male status. In response, females have evolved counter-strategies to reduce the impact of infanticide on their reproductive success. In conclusion and in consideration of the evidence available, both siblicide and infanticide life history strategies are adaptive even though they seem to be contradictory to the success of reproduction.

Common Criticisms in Psychology Paper

The artificiality of inventive conditions in experimental environments is a repeated concern. How real can laboratory-based research be? This paper will explain the criticism of artificiality in the discipline of psychology and apply this criticism to at least three sub disciplines within psychology. This paper will also compare and contrast the breakthrough model of scientific research and the principle of connectivity in explaining events and outcomes; finally ending with comparing and contrasting the concepts of the single cause explanation and the principle of multiple causation in explaining events and outcomes. Criticism of Artificiality Long gone are the days of William Wundt but what remains at the fore front is the expostulation of experimentation from critics that confining psychology to the laboratory spontaneously confines the mental phenomena it can analyze. An appropriate estimation of the artificiality criticism requires distinctively several intentions experimentalists follow. The discipline of psychology’s laboratory studies are seen by some as bizarre. Viewing psychology as an inadequate science by the public stems from belief that evidence cannot be acquired unless natural circumstances are examined (Stanovich, 2007). Social Psychology The college sophomore problem and criticisms of representativeness are most often aimed at social psychology, which makes frequent use of college subjects in laboratory paradigms in an attempt to develop theories of social interaction, group behavior, and information processing in social situations† ( as cited in Stanovich, 2007, page 114). Bio-medical research is correspondent in today’s state of social psychology, and many of times founded on problem searching and very well may be funded on the footing of the problem it intends to alleviate (Krueger, 2003). Popular and political interest plays a role in the search for a particular cure of disease. More over it is the methodical, theory based research of fundamental physiology that deciphers the operations of the human body’s functions as well as how under certain conditions it malfunctions (Fields, 1994; Skalka, 1993). In a correlating style, fundamental, â€Å"theory driven research on social psychological processes† will completely enlighten the distinctive defects and the adaptable successfulness of the social creature (Krueger, 2003). Abnormal Psychology In discussing the artificiality criticism within abnormal psychology, it has been shown that as a laboratory science, it has many times had to deal with solitary and maybe abnormal section of human behavior, more specifically college students; therefore inefficient of considering any ample warmth, richness, and inventiveness of human behavior. More concerns include the kind or representativeness of subjects participating in academic research. A quote taken from William Robinson (2007) sums up the artificiality criticism saying â€Å"If our interpretation of the human mind is based on the behavior of American college sophomores in artificially contrived situations and interpreted in terms of laws derived from domesticated rats and pigeons, then our psychology is truly culture bound† (Robinson, 2007). Organizational Psychology As the field of Organizational psychology has evolved and grown, so has the array of topics and important research questions needing answers. A common systematic approach implemented to start understanding work related psychological issues can be investigated. â€Å"Theory-driven and research-based human resource and organizational improvement efforts can be evaluated using state-of-the-art evaluation science approaches, and evaluation findings can be used to continually improve and institutionalize positive change efforts† ( Donaldson & Bligh, 2006). Connectivity and Convergence When a new theory in science touches upon previously verified empirical facts, this is definitive of the connectivity principle (Stanovich, 2007). Consideration as an advance is contingent upon explaining new facts while accounting for old ones. It makes no difference if the theory explains old facts differently from the earlier theory, just that they are explained. This necessity guarantees the accumulative development of science. Despite a new theory explaining new facts, if there is no explanation for the old ones then there are no considerations of finalized advancement and no immediate replacement of them (Stanovich, 2007). What will occur will be both the old and new theory coexisting as probable ideas until a new theory abdicates them null and void? Leading us astray is the breakthrough model of scientific research by suggesting that new findings violate the principle of connectivity (Stanovich, 2007). This suggestion deems risky because by abandoning the connectivity principle, the primary beneficiaries are purveyors of counterfeit science and incredulous theories. Notorious are headlines that lead off with â€Å"New Breakthrough. † Theories such as these acquire most of their interest and publicity by claims that they are astoundingly new. The next deception is to cast out past information by asserting them impertinent. The reason being given, that such information does not yet exist due to the newness of the theory. This breeds an environment of pseudoscience. Better explained is by the evolutionary theory the ghost of creationism, with its connectivity display of such different areas of science as morphology, biogeography, paleontology and many more. With the universe and earth estimating at around 10 thousand years in age, then many modern sciences of physics, chemistry, astronomy, cosmology, paleontology and beginning human history are entirely abolished (Unspoken Bible, n. . ). Darwin’s theory called pangenesis, abandons the principle of connectivity to illustrate the means heredity correlating with natural selection (Unspoken Bible, n. d. ). Where the problem lies is creationism proving no connectivity among any other things in science like in geology, genetics, ecology, chemistry and biology (Unspoken Bible, n. d. ). The utmost connectivity is proven with all the other sciences by evol ution (Unspoken Bible, n. d. ). Single Cause Explanation and Principle of Multiple Causation Even though a cause of behavior may be determined does not imply the only or most important cause of behavior (Stanovich, 2007). Causal analysis stems from the event explained as extreme, negative or unexpected. Preferences in single cause explanations may happen, such as when not enough time has been made looking for other possibilities, or a lack of cognitive resources. On the other hand, where constraints are lacking consideration in real effort may be given on a multitude of feasible causes for the event in question (Chu & Shaw, 2005). In providing a complete abdication of a specific behavior the weight of many different variables must be studied by the researcher as well as mixing the results of the studies in order to give a thorough snapshot of all the causal associations (Chu & Shaw, 2005). Despite an outcome having numerous different variable determinants this does not minimize the significance of a causally related outcome by one variable, this being the case for variables accounting for a mere percentage of the outcome. In considering multiple causes, explanations are often based on past experiences or intuitive theories (Chu & Shaw, 2005). This resulting in the accepted explanation that is most approachable or believable. Even though individuals understand and credit the existence of multiple causes they many times conduct themselves more along the lines of unitary beliefs (Stanovich, 2007). Complex behaviors are multiply determined. A multitude of factors perform to generate their occurrence. Mitigating a difference in the effect of acting together variables than what can be due easily studying them separately. Don’t fall prey to thinking that there is a single cause to a particular behavior. Conclusion In conclusion this paper has explained events and outcomes through compare and contrasts of both the breakthrough model of scientific research and the principle of connectivity and the concepts of single cause and the principle of multiple causation. A mutual consensus because psychology experiments are not like real life, this should be looked upon as a strength and not a weakness. No lone experiment is conclusive but instead gives leave of some alternative explanations, perpetuating the ability of zeroing in on the truth. References Chu, Y. & Shaw, J. (2005). Causal chaining: Effects of behavioral domain and Outcome valence on perceived causal structure. Retrieved August 10, 2009 from http://www. uiowa. edu/~grpproc/crisp/crisp10_14. pdf Donaldson, S. I. & Bligh, M. C. (2006). Rewarding careers applying positive psychological science to improve quality of work life and organizational effectiveness. In Donaldson, S. , Berger, D. & Pezdek, K. (Eds. ) Applied sychology: New frontiers and rewarding careers. ( pp. 277 – 295). Mahwah, NJ: Lawrence Erlbaum Associates. Fields, B. N. (1994). AIDS: Time to turn to basic science. Nature 369: 95 – 96. Krueger, J. (2003). Towards a balanced social psychology: Causes, consequences and cures fro the problem-seeking approach to social behavior and cognition. Retrieved August 10, 2009 from http://www. scribd. com Robinson, W. (2007). Is 531: Psychology. Retrieved August 10, 2009 from http://www. web. utk. edu Skalka, A. M. (1993). Lobbying for research dollars: as more money goes into specific diseases available for basic research diminish. Washington Post (June 22) WH6 Stanovich, K. (2007). How to think straight about psychology. (8th ed. ). Allyn & Bacon: Pearson Education Company. The Unspoken Bible. (n. d. ). The scientific method. Retrieved August 10, 2009 from http://www. usbible. com

Essay about South African Samples for Kids - a Brief Overview

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The Various Structures Of Empires And Global History

Utilizing three major events this essay will explore the relationship between the various structures of empires and global history. Understanding empires and imperial rules are fundamental understanding the reasons why and how certain events transpired. Beginning with a brief introduction of empires, including the general reasons empires expanded and how. Following the definition of empires, we will use the various social, political, and economic structures of empires to discuss how the slave trade, industrial revolution, and the scramble for Africa were a have a causal relationship with empires and imperial rule. Empires use political and economic power to expand into other territories and overtake the people by fostering†¦show more content†¦Socially, as before mentioned, they would create inequalities among the people native to that society. Politically, they would use military force to capture individuals and force them into slavery. Economically, an empire could impose a market on the colony restricting trade or they could use indirect control by making the state economically dependent on the Empire. Physical occupation was one means of expansion empires could impose their economic or political will creating the same dependent relationship between an empire and a colony (Class Notes, Feb 9). The strategy of dividing and conquering has been used from the earliest of empires. The Roman empire effectively used a strong army that conquered areas and took slaves for agricultural work. The Romans made strategic decisions about who were going to be slaves and those that would have limited rights (Laxer, p. 26 – 27), thus dividing the population and conquering vast amounts of land (Class Notes, Feb, 7). As with the Roman empire, the European empires understood that in order to expand the supply of labour needed to be tightly controlled; the development of the slave trade was a critical event that highlights how empires functioned and the resulting impact on global history. The Spanish empire’s search in Latin America was, not for the traditional resource of agriculture rather, for mineral reserves, such as gold and silver, wasShow MoreRelatedGlobalization Means Underaken Entirely805 Words   |  3 Pages1. Defining Globalization: The term Globalization in origin came from the word ‘global’. In different languages there are different meanings for the word global. Meydan Larousse defined the term global as ‘’undertaken entirely’’ and this meaning was attributed to the word global by western languages. In French this term means ‘’homogeneity’’. Henceforth the word means both â€Å"entirety† and â€Å"homogeneity†. 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